PDF] Comments on the Pteranodontidae (Pterosauria, Pterodactyloidea) with the description of two new species.

Based on several cranial features, some specimens previously referred to the genus Pteranodon are re-evaluated leading to the recognition of the following species, two of which new that are described here: PTeranodon longiceps, Geosternbergia sternbergi, Geosterbergia maiseyi sp. Considered one of the best known flying reptiles, Pteranodon has been subject to several reviews in the last century. Found exclusively in the Upper Cretaceous Niobrara Formation and Pierre Shale Group 11 species have been attributed to this genus (excluding the ones presently regarded as representing Nyctosaurus). While reviewers agree that this number is inflated, there is disagreement on how many species can be identified. The last review recognized only two species (Pteranodon longiceps and Pteranodon sternbergi) both being sexually dimorphic. Based on several cranial features, some specimens previously referred to the genus Pteranodon are re-evaluated leading to the recognition of the following species, two of which new that are described here: Pteranodon longiceps, Geosternbergia sternbergi, Geosternbergia maiseyi sp. nov., and Dawndraco kanzai gen. et sp. nov. They differ mainly by features such as the direction and extension of the frontal crest, the angle and extension of the posterior process of the premaxillae, the shape and extension of the lower temporal fenestra and the length and proportion of the rostrum. The procedures to recognize a pterosaur species are also discussed here, and must take into account primarily morphology, in conjunction with stratigraphic and geographic data. Although well aware that changes in morphology not always reflect taxonomy, the lack of stratigraphic data and the limited number of specimens that can be confidently assigned to one species hampers our understanding on the morphological variations as a function of ontogeny, individual variation and sexual dimorphism. Although the present study has not eliminated the possibility to recognize such differences, caution is needed before models are generalized for pterosaurs.

Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria)

New smallest specimen of the pterosaur Pteranodon and ontogenetic niches in pterosaurs, Journal of Paleontology

PDF) Comments on the Pteranodontidae (Pterosauria, Pterodactyloidea) with the description of two new species

Pteranodon sternbergi - Wikipedia

Pteranodon sternbergi - Wikipedia

November, 2011, The Pterosaur Heresies

PDF] Comments on Triassic pterosaurs with discussion about ontogeny and description of new taxa.

Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]

PDF] Comments on the Pteranodontidae (Pterosauria, Pterodactyloidea) with the description of two new species.

Dawndraco - Wikipedia

Full article: A new species of crested pterosaur (Pterodactyloidea, Anhangueridae) from the Lower Cretaceous (upper Albian) of Richmond, North West Queensland, Australia

New smallest specimen of the pterosaur Pteranodon and ontogenetic niches in pterosaurs, Journal of Paleontology

PDF) A New Crested Pterosaur from the Early Cretaceous of Spain: The First European Tapejarid (Pterodactyloidea: Azhdarchoidea)